by John P.T. Boorman,
6, Beckingham Road, Guildford, Surrey, GU2 6BN, United Kingdom
and Daniel V. Hagan,
Department of Biology, Institute of Arthropodology & Parasitology,
Georgia Southern University, Statesboro, GA 30460-8042 United States of
America
Key words: Diptera, Ceratopogonidae, Culicoides, name list,
taxonomy
Arnaud (1956) and Arnaud & Wirth (1964) published lists of World Culicoides up to 1962, listing a total of 916 names. Since that time, numerous new species have been described, but there has been no published work bringing this list up to date. Arnaud (1956) listed 671 names; Arnaud & Wirth added a further 245. The present paper includes all of these and names published up to 1994. It is hoped that the list will serve as a name check list and will serve to prevent homonymy. It has been distributed on disc partly because of its length, and also because users will thus be able to add to or amend their copy as they wish.
The presentation largely follows that of those quoted above; the names given are those proposed for species, subspecies or varieties within the genus Culicoides or which have been moved into that genus, but do not take account of synonymy and generic transfers subsequent to original publication. Nor has any account been taken of the subgeneric placement of the species; partly since the subgeneric classification of the genus is far from satisfactory, and partly because in the majority of cases such placement is open to doubt. The names are arranged alphabetically, followed by the name of the author and the year of publication. The journal title is given in full, following the practice now adopted by many major journals. The country or region from which the type originates is also given where possible, but the status and deposition of the type or syntypes has been omitted. This information can usually be had by consulting the publication concerned. Titles of papers have been omitted, for considerations of space. Finally, where a species has been moved into Culicoides, the name of the genus in which it was originally described is given in square brackets [ ].
The authors would be grateful if users would provide
notification of any mistakes, omissions or additions so that the list may
be kept up to date. Please send any corrections or changes to Dan
Hagan via the addresses above or e-mail, dhagan@GaSoU.edu.
abdominalis Santos Abreu, E. 1918. Essay for a monograph of the Tendipedidae of the Canary Islands (in Spanish). Memorias de la Real Academia de Ciencias y Artes de Barcelona, 14: 159-326. Canary Islands.
abreui Kieffer, J.J. 1921. Bulletin de la Societe Entomologique de France, 1921: 7 Canary Islands.
absitus Liu, C.W. & Yu, 1990. Contributions to Blood-Sucking Dipterous Insects, 2: 20. China.
acanthostomus Wirth, W.W. & A.A. Hubert. 1989. Memoirs of the American Entomological Institute, 44: 92. Malaysia.
*acastus de Meillon, B. 1947. New records and species of biting insects from the Ethiopian Region II. Journal of the Entomological Society of Southern Africa, 10: 110- 124. Transvaal.
*accraensis Carter, H.F., A. Ingram & J.W.S. Macfie. 1920. Observations on the Ceratopogonine midges of the Gold Coast with descriptions of new species pt. 2. Annals of Tropical Medicine & Parasitology, 14: 211-274. Ghana.
*achkamalicus Dzhafarov, S.M. 1964. Biting midges (Diptera, Heleidae) of Transcaucasus (Morphology, Biology, Ecology, Geographical Distribution, Harmfulness, Control and Fauna of the genera Culicoides, Leptoconops and Lasiohelea) [in Russian]. Akademia Nauk Azerbajdzhanskoi SSR Institut Zoologii, 263. Azerbajdzhan. 1964: 3-414.
*achrayi Kettle, D.S. & J.W.H. Lawson. 1955. Sex ratios among British Culicoides. Proceedings of the Royal Entomological Society of London (B), 30: 70-72. Scotland.
*acotylus Lutz, A. 1913. Contribution to the study of the bloodsucking Ceratopogonids of Brazil. Systematic Part. Second Memoir (in Portuguese and German). Memorias do Instituto Oswaldo Cruz, 5: 45-73, 3 plates. Brazil.
*actoni Smith, R.O.A. 1929. Two species of Culicoides which feed on man. Indian Journal of Medical Research, 17: 255-257, 1 plate. India.
adamskii Wirth, W.W. 1990. Proceedings of the Entomological Society of Washington, 92: 242. Aldabra.
*adersi Ingram, A. & J.W.S. Macfie. 1923. Notes on some African Ceratopogoninae. Bulletin of Entomological Research, 14: 41-74. Kenya.
*aethionotus Wirth, W.W. & F.S. Blanton. 1955. Studies in Panama Culicoides (Diptera, Heleidae). 5 descriptions of three new species of the subgenus Oecata Poey. Bulletin of the Brooklyn Entomological Society, 50: 121-127. Panama.
*africanus Clastrier, J. 1959. Notes on the Ceratopogonids. 6 Ceratopogonids of French West Africa (in French). Archives de l'Institut Pasteur d'Algerie, 37: 167-197. Senegal.
agas Wirth, W.W. & A.A. Hubert. 1989. Memoirs of the American Entomological Institute, 44: 438. Indonesia.
*agathensis Callot, J., M. Kremer & J.A. Rioux. 1963. On the Culicoides (Diptera: Ceratopogonids) with one species and one new variety from the Midi in France (in French). Annals de Parasitologie humaine et comparee, 38: 121-129. France.
*aitkeni Wirth, W.W. & F.S. Blanton. 1968. A revision of the neotropical biting midges of the Hylas group of Culicoides (Diptera, Ceratopogonidae). The Florida Entomologist, 51: 201-215. Trinidad.
*ajbassovi Shakirzhanova, M.S. 1962. New species of Culicoides (Diptera, Heleidae) from Kazakhstan (in Russian). Trudy Instituta Zoologii. Akademiya Nauk Kasakhskoi SSR, 18: 254-259. Kazakhstan.
*alachua Jamnback, H. & W.W. Wirth. 1963. The species of Culicoides related to obsoletus in Eastern North America (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 56: 185-198. Florida.
*alahialinus Barbosa, F.A.S. 1952. New additions to the knowledge of neotropical Culicoides (Diptera: Heleidae) (in Portuguese). University of Recife Thesis. Imprensa Industrial. Recife: 1952: 1-21, 10 plates. Ecuador.
*alambiculorum Macfie, J.W.S. 1948. Some species of Culicoides (Diptera, Ceratopogonidae) from the state of Chiapas, Mexico. Annals of Tropical Medicine & Parasitology, 42: 67-87. Mexico.
*alaskensis Wirth, W.W. 1951. The genus Culicoides in Alaska (Diptera, Heleidae). Annals of the Entomological Society of America, 44: 75-86. Alaska.
*alatavicus Smatov, Z.S. & N.K. Isimbekov. 1971. New and little known species of biting midges (Diptera, Ceratopogonidae) from Kazakhstan (in Russian). Izvestiya Akademii Nauk Kazakhskoi SSR, Trudy Instituta Zoologii 4:61-65. (n.n. for fuscus Gutsevich & Smatov, 1966)
*alatus Das Gupta, K.S. & S.M. Ghosh. 1956. On a new bloodsucking species Culicoides alatus, n. sp. (family Ceratopogonidae, order Diptera). Bulletin of the Calcutta School of Tropical Medicine, 4: 162-163. India.
*alazanicus Dzhafarov, S.M. 1961. Description of three species
of Heleidae from Asia Minor (in Russian). Izvestiya Akademii Nauk Azerbajdzhanskoi
SSR, Seriya Biologicheskikh i Meditinskikh Nauk no.10: 75-80. Azerbajdzhan.
*albertensis Wirth, W.W. & R.H. Jones. 1957. The North American
subspecies of Culicoides variipennis (Diptera, Heleidae). U.S. Department
of Agriculture Technical Bulletin no. 1170: 1-35. Alberta.
*albibasis Wirth, W.W. & A.A. Hubert. 1959. Trithecoides,
a new subgenus of Culicoides (Diptera, Ceratopogonidae). Pacific
Insects, 1: 1-38. Malaysia.
*albicans Winnertz, J. 1852. Contribution to the knowledge of
the genus Ceratopogon Meigen (in German). Linnaea Entomologica, 6: 1-80.
Germany [Ceratopogon]
*albidigaster Santos Abreu, E. 1918. Essay for a monograph of
the Tendipedidae of the Canary Islands (in Spanish). Memorias de la Real
Academia de Ciencias y Artes de Barcelona, 14: 159-326. Canary Islands.
*albidipes Santos Abreu, E. 1918. Essay for a monograph of the
Tendipedidae of the Canary Islands (in Spanish). Memorias de la Real Academia
de Ciencias y Artes de Barcelona, 14: 159-326. Canary Islands.
*albifascia Tokunaga, M. 1937. Sand flies (Ceratopogonidae, Diptera)
from Japan. Tenthredo, 1: 233-338, 6 plates. Taiwan.
*albihalter Kieffer, J.J. 1919. Chironomids of the Northern Polar
region (in German). Annales Historico-Naturales Musei Nationalis Hungarici,
8: 40-48, 111-120. Germany.
*albihalteratus Goetghebuer, M. 1935. Ceratopogonidae and Chironomidae
collected for the first time in Belgium (in French). Bulletin et Annales
de la Societe Royale Entomologique de Belgique, 75: 413-418. Belgium.
*albipennis Kieffer, J.J. 1919. Chironomids from Europe preserved
in the National Hungarian Museum of Budapest (in French). Annales Historico-Naturales
Musei Nationalis Hungarici, 17: 1-160. France.
*albipennis Smith, R.O.A., & C.S. Swaminath. 1932. Notes
on some Culicoides from Assam. Indian Medical Research Memoires,
25: 182-186, 1 plate. Assam.
*alboguttatus Kieffer, J.J. 1921. Chironomids from the Philippines
and Formosa (in French). Philippine Journal of Science, 18: 557-593. Taiwan.
*albohalteratus Santos Abreu, E. 1918. Essay for a monograph
of the Tendipedidae of the Canary Islands (in Spanish). Memorias de la
Real Academia de Ciencias y Artes de Barcelona, 14(2): 159-326. Canary
Islands.
*albomacula Root, F.M., & W.A. Hoffman. 1937. The North American
species of
Culicoides. American Journal of Hygiene, 25: 150-176,
8 plates. Mexico.
*albonotatus Kieffer, J.J. 1918. Chironomids from Africa and
Asia preserved in The National Hungarian Museum of Budapest (in French).
Annales Historico-Naturales Musei Nationalis Hungarici, 16: 31-136. Asia
Minor.
*albonotatus Vimmer, A. 1932. New species of the subfamily Ceratopogoninae
(Tendipedidae, Dipt.) from the Bodenheimer Collection (in Czech). Sbornik
Entomologickeho Oddeleni Narodniho Musea v Praze, 10: 130-144. Palestine.
*albopunctatus Clastrier, J. 1960. Notes on the Ceratopogonids.
10 Ceratopogonids of the Republic of the Congo (2) (in French). Archives
de l'Institut Pasteur d'Algerie, 38: 258-298. Congo Republic.
*albosignatus Vimmer, A. 1932. New species of the subfamily Ceratopogoninae
(Tendipedidae, Dipt.) from the Bodenheimer Collection (in Czech). Sbornik
Entomologickeho Oddeleni Narodniho Musea v Praze, 10: 130-144. Palestine.
*albosparsus Kieffer, J.J. 1918. Chironomids from Africa and
Asia preserved in The National Hungarian Museum of Budapest (in French).
Annales Historico-Naturales Musei Nationalis Hungarici, 16: 31-136. Ethiopia.
*albovenosus Khamala, C.M.P., & D.S. Kettle. 1971. The Culicoides
Latreille (Diptera: Ceratopogonidae) of East Africa. Transactions of the
Royal Entomological Society of London, 123: 1-95. Uganda.
*albuquerquei Wirth, W.W. & F.S. Blanton. 1973. A review
of the maruins of biting midges of the genus Culicoides (Diptera:
Ceratopogonidae) in the Amazon Basin. Amazoniana, 4: 405-470. Brazil.
*alexanderi Wirth, W.W., & A.A. Hubert. 1962. The species
of Culicoides related to Piliferus and Hoffman in Eastern North
America (Diptera: Ceratopogonidae). Annals of the Entomological Society
of America, 55: 182-195. Massachusetts.
*alexandrae Dzhafarov, S.M. 1962. New species of bloodsucking
midges (Diptera, Heleidae) from the valley of the Kura River, Transcaucasus
(in Russian). Entomologicheskoe Obozrenie, 41: 241-246. Azerbajdzhan.
*alexis Meillon, B. De. 1936. Entomological studies. Studies
on insects of medical important from South Africa. Part III. South African
Ceratopogonids. Part I. The species recorded from South Africa. Publications
of the South African Institute for Medical Research, 7: 136-140. South
Africa.
*algarum Kieffer, J.J. 1924. New or rare Chironomids from Central
Europe (in French). Bulletin de la societe d'Histoire Naturelle de la Moselle,
30: 11-110. Germany.
*algecirensis Strobl, G. 1900. Spanish Diptera. Part XI. (in
German). Wiener Entomologische Zeitung, 19: 169-174. Spain. [Ceratopogon]
*algeriensis Clastrier, J. 1957. Notes on the Ceratopogonids.
2. Some Culicoides from Algeria with spotted wings (in French).
Archives de l'Institu Pasteur d'Algerie, 35: 404-444. Algeria.
alishanensis Chen, 1988. Chinese Journal of Entomology, 8: 151.
Taiwan.
allantothecus Wirth, W.W., & A.A. Hubert. 1989. Memoirs of
the American Entomological Institute, 44: 121. Malaysia.
*almeidae Cambournac, F.J.C. 1970. Culicoides almeidae
(Nematocera, Ceratopogonidae) Sp. N. A new species encountered in Portugal
(in Portuguese). Anais da Escola National de Saude Publica e de Medicina
Tropical, 4: 251-257. Portugal.
*almirantei Wirth, W.W., & F.S. Blanton. 1959. Biting midges
of the genus Culicoides from Panama (Diptera: Heleidae). Proceedings
of the U.S. National Museum, 109: 237-482. Panama.
*altaicus Remm, H. 1972. New species of Ceratopogonidae (Diptera)
from the South Siberia. (in Russian, English Summary). Tartu Riikliku Ulikooli
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*alticola Kieffer, J.J. 1913. Chironomidae and Cecidomyidae (in
French). Voyage de Ch. Alluaud et R. Jeannel en Afrique orientale, 1911-1912:
1-43. Africa.
*alvarezi Ortiz, I. 1957. New bloodsucking midges representatives
of the genera Culicoides (Diptera: Simuliidae) from Venezuela (in
Spanish). Boletin Venezolano del Lab. Clin., 2: 161-168. Venezuela.
*amamiensis Tokunaga, M. 1937. Sand flies (Ceratopogonidae, Diptera)
from Japan. Tenthredo, 1: 233-338, 6 plates. Ryukyu Islands.
*amaniensis Khamala, C.P.M., & D.S. Kettle. 1971. The Culicoides
Latreille (Diptera: Ceratopogonidae) of East Africa. Transactions of the
Royal Entomological Society of London, 123: 1-95. Tanzania.
*amazonius Macfie, J.W.S. 1935. Ceratopogonidae (Dipt.) from
the River Amazon. Stylops, 4: 49-56. Brazil.
ameliae Browne, J.G. 1980. Journal of Medical Entomology, 17:
543. Colombia.
*amoenus Winnertz, J. 1852. Contribution to the knowledge of
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*amossovae Remm, H. 1971. On the fauna of Ceratopogonidae of
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anadyriensis Mirzaeva, A.G. 1984. Chlenistonogie i Gel'minty
Novye i Maloizvestnye Vidy Fauny Sibiri, vyp. 17: 70. USSR.
*analis Santos Abreu, E. 1918. Essay for a monograph of the Tendipedidae
of the Canary Islands (in Spanish). Memorias de la Real Academia de Ciencias
y Artes de Barcelona, 14(2): 159-326. Canary Islands.
*analis Kieffer, J.J. 1925. Diptera (biting Nematocera): Chironomidae
Ceratopogoninae (in French). Faune de France, 11: 1-139. Belgium.
*andicola Wirth, W.W., & V.H. Lee. 1967. New species of Culicoides
from high altitudes in the Colombian Andes (Diptera: Ceratopogonidae).
Proceedings of the U.S. National Museum, 124: 1-22. Colombia.
*andinus Wirth, W.W., & V.H. Lee. 1967. New species of Culicoides
from high altitudes in the Colombian Andes (Diptera: Ceratopogonidae).
Proceedings of the U.S. National Museum, 124: 1-22. Colombia.
*andrewsi Causey, O.R. 1938. Culicoides of Siam with descriptions
of new species. American Journal of Hygiene, 27: 399-416, 8 plates. Thailand.
*angolensis Caeiro, V.M.P. 1961. Contribution to a study of the
Angolan species of the genus Culicoides Latreille 1809 (in Portuguese).
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*angularis Lee, D.J., & E.J. Reye. 1953. Australasian Ceratopogonidae
(Diptera, Nematocera) Part 6. Australian species of Culicoides.
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annuliductus Wirth, W.W. 1981. Proceedings of the Entomological
Society of Washington, 83: 150. Panama.
*anophelis Edwards, F.W. 1922. On some Malayan and other species
of Culicoides, with a note on the genus Lasiohelea. Bulletin of
Entomological Research, 13: 161-167. Malaysia, India.
*antefurcatus Wirth, W.W., & F.S. Blanton. 1959. Biting midges
of the genus Culicoides from Panama (Diptera: Heleidae). Proceedings
of the U.S. National Museum, 109: 237-482. Panama.
*antennalis Lee, D.J., & E.J. Reye. 1953. Australasian Ceratopogonidae
(Diptera, Nematocera) Part 6. Australian species of Culicoides.
Proceedings of the Linnaean Society of New South Wales, 77: 369-394. Australia.
aomoriensis Kitaoka, 1991. Japanese Journal of Sanitary Zoology,
42: 289. Japan.
apiculatus Yu & Zhang, 1988. China Public Health (suppl.),
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*aquilinus Smatov, Z.S., & G.A. Kravets. 1976. New species
of midges (Diptera, Ceratopogonidae, Culicoides) from Kazakstan
(in Russian, English Summary). Parazitologiya, 10: 282-286. Kazakhstan.
arabiensis Boorman, 1989. Fauna of Saudi Arabia, 10: 173. Oman.
aragaoi Tavares & Luna Dias, 1980. Revista Brasileira de
Biologia, 40: 393. Brazil.
*arakawae Arakawa, S. 1910. On a new injurious "Insect" (Ceratopogon
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(as arakanae)
*arboreus Gutsevich, A.V. 1952. Contribution to the fauna of
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*arboricola Root, F.M., & W.A. Hoffman. 1937. The North American
species of Culicoides. American Journal of Hygiene, 25: 150-176,
8 plates. Maryland.
*archboldi Wirth, W.W., & F.S. Blanton. 1970. New genera
of neotropical Culicoides (Diptera: Ceratopogonidae). The Florida
Entomologist, 53: 7-14. Dominica.
*arcuatus Winnertz, J. 1852. Contribution to the knowledge of
the genus Ceratopogon Meigen (in German). Linnaea Entomologica, 6: 1-80.
Europe.
*ardentissimus Tokunaga, M. 1940. Ceratopogonidae and Chironomidae
from the Micronesian Islands with biological notes by Teiso Esaki. Philippine
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*ardleyi Tokunaga, M. 1962. Biting midges of the genus Culicoides
from the New Guinea (Diptera: Ceratopogonidae). Pacific Insects, 4: 457-516.
New Guinea.
*arenarius Edwards, F.W. 1922. On some Malayan and other species
of Culicoides, with a note on the genus Lasiohelea. Bulletin of
Entomological Research, 13: 161-167. Somalia.
arenicola Howarth, 1985. International Journal of Entomology,
27: 79. Laos.
*aricola Kieffer, J.J. 1922. Biological observations on the biting
Chironomids with a description of two new species (in French). Archives
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of Okinawa and the description of two new species. Proceedings of the Entomological
Society of Washington, 63: 235-239. Ryukyu Islands.
arschanicus Mirzaeva, 1984. Chlenistonogie i Gel'minty Novye
i Maloizvestnye Vidy Fauny Sibiri, vyp. 17: 69. USSR.
*arubae Fox, I., & W.A. Hoffman. 1944. New neotropical biting
sand flies of the genus
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*assamensis Smith, R.O.A., & C.S. Swaminath. 1932. Notes
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*assimilis Delfinado, M.D. 1961. The Philippine biting midges
of the genus Culicoides (Diptera: Ceratopogonidae). Fieldiana: Zoology,
33: 627-675, 7 plates. Philippine Islands.
*atchleyi Wirth, W.W., & F.S. Blanton. 1969. North American
Culicoides
of the Pulicaris group (Diptera: Ceratopogonidae). Proceedings of the Entomological
Society of Washington, 52: 207-243. Alaska.
atelis Wirth, 1982. The Florida Entomologist, 65: 249. Panama.
*aterinervis Tokunaga, M. 1937. Sand flies (Ceratopogonidae,
Diptera) from Japan. Tenthredo, 1: 233-338, 6 plates. Japan.
*atripalpis Wirth, W.W., & F.S. Blanton. 1973. A review of
the Maruins or biting midges of the genus Culicoides (Diptera: Ceratopogonidae)
in the Amazon Basin. Amazoniana, 4: 405-470. Brazil.
*atripennis Shevchenko, A.K. 1972. Description of two new species
of blood-sucking biting midges of the genus Culicoides (Diptera,
Ceratopogonidae) from the Ukraine (in Russian, English Summary). Vestnik
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*aurantiacus Kieffer, J.J. 1911. The Percy Siaden Trust Expedition
to the Indian Ocean in 1905 under the leadership of Mr. J. Stanley Gardiner,
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Society, 14: 331-366, 1 plate. Seychelles.
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(Ortiz, 1950). Description of four new species and the redescription of
some others less known (in Spanish). Revista de Sanidad y Asistancia Social,
16: 573-591, 8 plates. Venezuela.
*austeni Carter, H.F.A., Ingram, & W.S. Macfie. 1920. Observations
on the Ceratopogonine midges of the Gold Coast with descriptions of new
species Pt. 2. Annals of Tropical Medicine & Parasitology, 14: 211-274,
2 plates. Ghana.
*australiensis Kieffer, J.J. 1917. Chironomids from Australia
preserved in the National Hungarian Museum of Budapest (in French). Annales
Historico-Naturales Musei Nationalis Hungarici, 15: 175-228. New Guinea.
*australis Wirth, W.W., & R.H. Jones. 1957. The North American
subspecies of Culicoides variipennis (Diptera, Heleidae). U.S. Department
of Agriculture Technical Bulletin no.1170: 1-35. Louisiana.
*austropalpalis Lee, D.J., & E.J. Reye. 1954. Australasian
Ceratopogonidae (Diptera, Nematocera). Part VII. Notes on the genera Alluaudomyia,
Ceratopogon, Culicoides and Lasiohelea. Proceedings of the Linnaean
Society of New South Wales, 79: 233-246. (n.n. for palpalis Lee & Reye)
*autumnalis Sen, P., & S.K. Das Gupta. 1959. Studies on Indian
Culicoides
(Ceratopogonidae: Diptera). Annals of the Entomological society of America,
52: 617-630. India.
*avilaensis Ortiz, I., & M. Mirsa. 1951. Studies on Culicoides.
Description of two new species: Culicoides avilaensis and C.
discrepans and the male of C. leopoldoi Ortiz, 1951. Redescription
of C. limai Barretto, 1944, C. baueri Hoffman, 1925, C.
lichyi Floch and Abonnenc and C. pusillus Lutz, 1913 (in Spanish).
Revista de Sanidad y Asistencia Social, 16: 593-605, 5 plates. Venezuela.
*azerbajdzhanicus Dzhafarov, S.M. 1962. New species of bloodsucking
midges (Diptera, Heleidae) from the valley of the Kura River, Transcaucasus
(in Russian). Entomologicheskoe Obozrenie, 41: 206-219 (Translation in
Ent. Rev. 41: 121-131). Azerbajdzhan.
*azureus Wirth, W.W., & F.S. Blanton. 1959. Biting midges
of the genus Culicoides from Panama (Diptera: Heleidae). Proceedings
of the U.S. National Museum, 109: 237-482. Panama.
*babrius de Meillon, B. 1943. New records, and new species of
Nematocera (Diptera) from the Ethiopian Region. Journal of the Entomological
Society of Southern Africa, 6: 90-113. Zimbabwe.
*badooshensis Khalaf, K.T. 1961. More Culicoides from
Iraq (Diptera: Heleidae). Beitrage zur Entomologie, 11: 450-471. Iraq.
*baghdadensis Khalaf, K.T. 1957. Heleids from Iraq, with description
of new species (Diptera: Heleidae (Ceratopogoninae)). Bulletin de la Societe
Entomologique d'Egypte, 41: 335-350. Iraq.
*bahiensis Barbosa, F.A.S. 1947. Culicoides (Diptera:
Heleidae) of the neotropical region (in Portuguese). Anais da Sociedade
de Biologia de Pernambuco, 7: 3-30, 9 plates. Brazil.
bahrainensis Boorman, 1989. Fauna of Saudi Arabia, 10: 177. Bahrain.
*baisasi Wirth, W.W., & A.A. Hubert. 1959. Trithecoides,
a new subgenus of Culicoides (Diptera, Ceratopogonidae). Pacific
Insects, 1: 1-38. Philippine Islands.
bajensis Wirth, W.W., & Moraes, 1979. Pan-Pacific Entomologist,
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*bakeri Vargas, L. 1954. Two new species of Mexican Culicoides
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