by Dr. Doug Kettle, Emeritus Professor
Department of Entomology, University of Queensland,
Queensland 4072, Australia
on November 23^rd 1994

* Terry Linley (John's widow) asked to make some additions to Dr. Kettle's obituary for John. Terry's additions are set off in italics from Dr. Kettle's remarks.

    I was deeply moved to learn of the death of John Linley on November 19th. He had always appeared to be supremely fit so that I had thought that one day he might write my obituary, but I never expected to be composing one for him. I first met John in the middle of 1959, when he had completed a B.Sc. degree at the University College of the West Indies, in special relationship with the University of London, and he joined me as a research assistant on the newly created Sandfly Research Unit in Montego Bay, Jamaica. A photograph of young John and the other members of the team can be found in the Bulletin of Entomological Research (1968), 58. plate XXX, following  p. 753.

    Shortly after joining the unit he celebrated his 21st birthday and quickly became one of the family, being much admired by my young children. He carried the aura of being the 200 m butterfly swimming champion of the Caribbean and he was the only person I have ever known to emerge sweating after a training session in sea or pool! John, my wife and I sang in a choir which performed on formal occasions on request and competed with some success in the local choral competitions. Another cameo of John is of him flapping the front of his shirt to cool off.

    Towards the end of my two year secondment to the Govt. of Jamaica a new sandfly problem emerged in the tourist area, caused by Leptoconops bequaerti, which had been little studied in the past. John immediately began a detailed study of that species and I left him happily engaged on that project when I returned to Edinburgh, Scotland at the end of 1960. A couple of years later he went to the Western Reserve University in Cleveland, Ohio where he met and married Sue. They returned shortly afterwards to Jamaica and John rejoined the Sandfly Research Unit.

    Following the Zoological Congress in Washington in 1963 I flew down to Jamaica to see how the work was progressing and stayed with John and Sue who were ecstatically happy at the birth of their daughter in May of that year. John and Sue had their second daughter Lori in 1964.

    In 1965 I caught up with John again when he was working at the London School of Hygiene and Tropical Medicine and I was on study leave from the University of Nairobi, Kenya. John was not suited to the life of a commuter travelling 25 miles from Guildford to Central London in the morning and back again in the evening. As a medical establishment there was a formality in dress at the L.S.H.T.M. which John found difficult to accept. Eccentricity was acceptable in professorial staff but was definitely frowned upon in the lower ranks.

    John had the disappointment of having his Ph.D. thesis returned for revision and resubmission. I thought that the request that he resubmit was not entirely reasonable, especially when I learned the name of the dissenting examiner. He/She was voicing a complaint which had been levelled against the examiner so perhaps he/she was ultrasensitive on the issue. Nevertheless John disciplined himself to revise and resubmit the thesis, which was accepted without comment. After arriving in Vero Beach in 1967 to take a job at the then Entomological Research Center, their third daughter Diane was born. From 1969 to 1985 John was married to Patricia Linley.

    John showed his quality as a scientist by later submitting a thesis for the highest degree awarded by the University of London, a Doctor of Science (D.Sc.). That degree is awarded on published work which develops a theme within a discipline and which has been published long enough for informed criticism to have been made. When joint papers are included in such a thesis co-authors are required to present signed statements on the contributions made by the individual authors. John well deserved the award of the D.Sc. (Lond.). When the Florida Medical Entomology Laboratory became associated with the University of Florida and John's status was confirmed by him being made a full professor of the University.

    We next met at the International Congress of Entomology in Canberra, Australia in 1972 when John accompanied my wife and me back to Brisbane where I was able to show him some of the State's beauty and also the breeding sites of coastal Culicoides and Leptoconops on Bribie Island. It was not until 1980 that we met in person again in London in 1980 at the Fourth International Symposium on Ceratopogonidae, a three day meeting held in the British Museum (Natural History), London and the Animal Virus Research Institute at Pirbright (See Israel Journal of Entomology (1981) 15, 109-128.

    In 1984 I was able to spend some days at Vero Beach on my way to the International Congress of Tropical Medicine and Malaria in Calgary and share with John his enthusiasm for his current activities and the application of new techniques as well as appreciating his past achievements. We were able to visit breeding sites of blood-sucking ceratopogonids on the Indian River, one of which is appropriately named Leptoconops linleyi. John appreciated having Bill Wirth, the doyen of ceratopogonid taxonomists, near at hand in Gainesville when Bill moved south after retirement. In the last three months a fruitful era in the study of ceratopogonids has come to an end with the deaths of Bill Wirth and John Linley.

    In 1986 John married Terry Reed; his fourth daughter Erin was born in February 1987 and his only son Brian Roger was born in July 1989.

    John's last effort was to convene the Ceratopogonid Symposium during the Dipterology Congress in Guelph in August this year. It required much patience and tact to make it possible for Dr. Valentina Glukhova to attend the Congress from the Zoological Institute in St. Petersburg and participate in the symposium. It was entirely due to John's persistence that she was able to attend.

    In a letter earlier this year John referred to his intention to take his family to visit Lisa and to see his first grandson and then go to England in September to bring his father over to the States to be close to him and his brother, Martin, who was due to move to Vero Beach from Jamaica in July. It is an irony of life that having brought the family physically close together so that his father might have the benefit of his sons to care for him in his later years that John should die first. I find it difficult to believe that John is dead. To me it is the death of an admired and much loved younger member of the family.

* Terry Linley (John's widow) asked to make some additions to Dr. Kettle's obituary for John. Terry's additions are set off in italics from the Dr. Kettle's remarks. Terry was very grateful that Dr. Kettle shared his memories of John with the group, and she wanted to expand a little bit on some details of John's life.

Linley, J.R. 1964. A description of the larvae and pupae of Culicoides furens Poey and Culicoides hoffmani Fox (Diptera: Ceratopogonidae). Ann. Mag. Nat. History 7: 129-149.

Linley, J.R. 1965. Autogeny in Jamaican sandflies (Ceratopogonidae). Proc. XIII Int. Congr. Ent. 800-801.

Linley, J.R. 1965. Descriptions of the pupae of Culicoides barbosai, C. insignis, C. borinqueni (Diptera: Ceratopogonidae). Ann. Ent. Soc. Amer. 58: 57-63.

Linley, J.R. 1965. Changes in the ovaries of certain biting midges (Diptera: Ceratopogonidae) following completion of the gonotrophic cycle. Mosquito News 25: 306-310.

Linley, J.R. 1965. The ovarian cycle and egg stage in Leptoconops (Holoconops) becquaerti (Kieff.) (Diptera: Ceratopogonidae). Bull. Ent. Res 56: 37-56.

Linley, J.R. 1965. Techniques for obtaining viable eggs of Leptoconops becquaerti Kieffer, Culicoides furens Poey and Culicoides barbosai Wirth and Blanton (Diptera: Ceratopogonidae). Mosquito News 25: 452-456.

Linley, J.R. and Davies, J.B. 1965. Breeding sites of Leptoconops bequaerti Kieffer in Jamaica. Trans. R. Soc. Trop. Med. Hyg. 59: 362-363.

Davies, J.B. and Linley, J.R. 1965. Observations on the breeding sites of the sandfly Leptoconops becquaerti in the Montego Bay area of Jamaica, with a note on one breeding site in Honduras. Caribbean J. Sci. 5: 117-128.

Linley, J.R. 1966. Field and laboratory observations on the behavior of the immature stages of Culicoides furens Poey (Diptera: Ceratopogonidae). J. Med. Ent. 2: 385-391.

Davies, J.B. and Linley, J.R. 1965. A standard flotation method for separating Leptoconops (Diptera: Ceratopogonidae) and other larvae from sand samples. Mosquito News 26: 440.

Linley, J.R. 1966. Rearing larvae of Leptoconops bequaerti Kieffer (Diptera: Ceratopogonidae) in the field. Trans. R. Soc. Trop. Med. Hyg. 60: 21.

Linley, J.R. 1966. The ovarian cycle in Culicoides barbosai Wirth and Blanton and C. furens (Poey) (Diptera: Ceratopogonidae). Bull. Ent. Res. 57: 1-17.

Linley, J.R. 1966. Effects of supplementary carbohydrate feeding on fecundity and life-length in Leptoconops becquaerti (Kieff.). Bull. Ent. Res. 57: 19-52.

Kettle, D.S. and Linley, J.R. 1967. The biting habits of Leptoconops becquaerti. I. Methods; standardization of technique; preferences for limbs, individuals and positions. J. Appl. Ecol. 4: 379-395.

Kettle, D.S. and Linley, J.R. 1967. The biting habits of Leptoconops becquaerti. II. Effect of meteorological conditions on biting activity; 24 hours and seasonal cycles. J. Appl. Ecol. 4: 397-420.

Linley, J.R. 1968. Autogeny and polymorphism for wing length in Leptoconops becquaerti (Kieff.) (Diptera: Ceratopogonidae). J. Med. Ent. 5: 53-66.

Linley, J.R. and Evans, F.D.S. 1968. A simple device to aid time-lapse studies with insects. Ann. Ent. Soc. Amer. 61: 775-777.

Linley, J.R. 1968. Studies on the larval biology of Leptoconops becquaerti (Kieff.) (Diptera: Ceratopogonidae). Bull. Ent. Res. 58: 1-24.

Linley, J.R. 1968. Colonization of Culicoides furens. Ann. Ent. Soc. Amer. 61: 1486-1490.

Linley, J.R. and Nielsen, H.T. 1968. Transmission of a mosquito iridescent virus Aedes taeniorhynchus. I. Laboratory experiments. J. Invert. Path. 12: 7-16.

Linley, J.R. and Nielsen, H.T. 1968. Transmission of a mosquito iridescent virus Aedes taeniorhynchus. II. Experiments related to transmission in nature. J. Invert. Path. 12:17-24.

Kettle, D.S. and Linley, J.R. 1969. The biting habits of some Jamaican Culicoides. I. C. barbosai Wirth and Blanton. Bull. Ent. Res. 58: 729-753.

Linley, J.R. 1969. Studies on larval development in Culicoides furens (Poey) (Diptera: Ceratopogonidae). I. Establishment of a standard rearing technique. Ann. Ent. Soc. Amer. 62: 702-711.

Linley, J.R. 1969. Seasonal changes in larval populations of Leptoconops becquaerti (Kieff.) (Dipt., Ceratopogonidae)in Jamaica, with observations on the ecology. Bull. Ent. Res. 59: 47-64.

Kettle, D.S. and Linley, J.R. 1969. The biting habits of some Jamaican Culicoides. II. C. furens (Poey). Bull. Ent. Res. 59: 1-20.

Linley, J.R., Evans, H.T. and Evans, F.D.S. 1970. A quantitative study of autogeny in a naturally occurring population of Culicoides furens (Poey) (Diptera: Ceratopogonidae). J. Anim. Ecol. 39: 169-183.

Linley, J.R. 1970. Description of the pupa of Culicoides floridensis. Ann. Ent. Soc. Amer. 63: 1016-1019.

Linley, J.R. 1970. The immature stages of Culicoides arboricola Root and Hoffman (Diptera: Ceratopogonidae). J. Med. Ent. 7: 717-721.

Linley, J.R., Evans, F.D.S. and Evans, H.T. 1970. Seasonal emergence of Culicoides furens (Diptera: Ceratopogonidae) at Vero Beach, Florida. Ann. Ent. Soc. Amer. 63: 1332-1339.

Linley, J.R. and Davies, J.B. 1971. Sandflies and tourism in Florida and the Bahamas and Caribbean area. J. Econ. Ent. 64: 264-278.

Linley, J.R. and Adams, G. M. 1971. Form and function of the spermatophore in Culicoides melleus (Coq.) (Dipt., Ceratopogonidae). Bull. Ent. Res. 60: 427-430.

Linley, J.R. and Evans, D.G. 1971. Behavior of Aedes taeniorhynchus larvae and pupae in a temperate gradient. Ent. Exp. Appl. 14: 319-332.

Linley, J.R. and Adams, G.M. 1972. Ecology and behaviour of immature Culicoides melleus (Coq.) (Dipt., Ceratopogonidae). Bull. Ent. Res. 62: 113-127.

Linley, J.R. and Adams, G.M. 1972. A study of the mating behaviour Culicoides melleus (Coquillett) (Diptera: Ceratopogonidae). Trans. R. Ent. Soc. Lond. 124: 81-121.

Hinds, M.J. and Linley, J.R. 1974. Changes in male potency with age after emergence in the fly Culicoides melleus. J. Insect Physiol. 20: 1037-1040.

Linley, J.R. and Cheng, L. 1974. The grooming organs of Culicoides (Diptera: Ceratopogonidae). Mosquito News 34: 204-206.

Linley, J.R. and Hinds, M.J. 1974. Male potency in Culicoides melleus (Coq.) (Diptera, Ceratopogonidae). Bull. Ent. Res. 64: 123-128.

Linley, J.R. and Adams, G.M. 1975. Sexual receptivity in Culicoides melleus (Diptera: Ceratopogonidae). Trans. R. Ent. Soc. Lond. 126: 279-303.

Linley, J.R. and Hinds, M.J. 1975. Quantity of the male ejaculate influenced by female unreceptivity in the fly, Culicoides melleus. J. Insect Physiol. 21: 281-285.

Linley, J.R. 1975. Termination of copulation and associated behaviour in Culicoides melleus (Coq.) (Diptera: Ceratopogonidae). Bull. Ent. Res. 65: 143-156.

Linley, J.R. and Hinds, M.J. 1975. Sperm loss at copulation of Culicoides melleus (Coq.) (Diptera: Ceratopogonidae). J. Ent. (A) 50: 37-41.

Linley, J.R. and Mook, M.S. 1975. Behavioural interaction between sexually experienced Culicoides melleus (Coquillett) (Diptera: Ceratopogonidae). Behaviour 44: 97-110.

Linley, J. R. 1975. Sperm supply and its utilization in doubly inseminated flies, Culicoides melleus. J. Insect Physiol. 21: 1785-1788.

Linley, J.R. 1976. Biting midges of mangrove swamps and salt-marshes (Diptera: Ceratopogonidae). In 'Marine Insects', Ed. L. Cheng. North Holland publishing Co. 335-376.

Linley, J.R. and Hinds, M.J. 1976. Seasonal changes in size, female fecundity and male potency in Culicoides melleus (Diptera: Ceratopogonidae). J. Med. Ent. 13: 151-156.

Linley, J.R. 1977. The role of the female in the sexual behavior of Culicoides melleus. Mosquito News 37: 283.

Linley, J.R. and Hinds, M.J. 1977. Ejaculation and spermatophore formation in Culicoides melleus. Mosquito News 37: 283-284.

Lowrie, R.C., Orihel, T.C. and Linley, J.R. 1977. Neotropical filariasis: vector studies involving Culicoides. Mosquito News 37: 286-287.

Linley, J.R. and Carlson, D.A. 1978. A contact mating pheromone in the biting midge, Culicoides melleus. J. Insect Physiol. 24: 423-427.

Linley, J.R. and Mook, M.S. 1979. Seasonal change in the pupal sex ratio in a population of Culicoides melleus (Coq.) (Diptera: Ceratopogonidae). Proc. Fla. Anti-Mosquito Assn. 49:6-11.

Linley, J.R. 1979. Nutrient agar mixtures for culture of food organisms for predaceous Culicoides larvae (Diptera: Ceratopogonidae). J. Med. Ent. 16: 171-172.

Linley, J.R. 1979. Activity and motility of spermatozoa of Culicoides melleus (Coquillett)(Diptera: Ceratopogonidae). Ent. Exp. Appl. 26: 85-96.

Linley, J.R. 1979. Some potential food organisms for rearing predaceous Culicoides larvae. Annis. Parasit. Hum. Comp. 54: 253-254.

Linley, J.R. 1979. Some aspects of sperm transfer in Culicoides melleus. Annis. Parasit. Hum. Comp. 54: 254-255.

Linley, J.R. 1980. Laboratory colonization of Ceratopogonidae (Diptera)-problems and solutions. Proc. 3rd European Multicolloquium of Parasitology. 93 p.

Linley, J.R. 1981. Ejaculation and spermatophore formation in Culicoides melleus (Coq.) (Diptera: Ceratopogonidae). Can. J. Zool. 59: 332-346.

Linley, J.R. 1981. Emptying of the spermatophore and spermathecal filling in Culicoides melleus (Coq.)(Diptera: Ceratopogonidae). Can. J. Zool. 59: 347-356.

Linley, J.R. and Simmons, K.R. 1981. Sperm motility and spermathecal filling in lower Diptera. Internat. J. Invert. Reprod. 4: 137-146.

Linley, J.R. and Simmons, K.R. 1983. Quantitative aspects of sperm transfer in Simulium decorum Walker (Diptera: Simuliidae). J. Insect Physiol. 29: 581-584.

Linley, J.R. 1982. Precocious autogeny in Leptoconops linleyi Wirth and Atchley. Proc. Fla. Anti-Mosquito Assn. 53: 1-3.

Linley, J.R. and Carlson, D.A. 1983. The effect of age, freezing and wing removal on the sexual response elicited by female Culicoides melleus (Diptera: Ceratopogonidae). J. Med. Ent. 20: 81-86.

Linley, J.R., Hoch, A.L. and Pinheiro, F.P. 1983. Biting midges (Diptera: Ceratopogonidae) and human health. J. Med. Ent. 20: 347-364.

Linley, J.R. 1983. Autogeny in the Ceratopogonidae: literature and notes. Fla. Ent. 66: 228-234.

Linley, J.R. 1983. Bodily stimulus gradients and precopulatory orientation in the midge, Culicoides melleus. Physiol. Ent. 8: 403-412.

Linley, J.R. and Braverman, Y. 1984. The tergal pigmentation patterns of Culicoides variipennis (Coq.) and Culicoides furens (Poey) (Diptera: Ceratopogonidae). J. Med. Ent. 21: 636-647.

Linley, J.R. 1984. The timing of diuresis in newly emerged adults of Culicoides variipennis (Coq.) (Diptera: Ceratopogonidae). Mosquito News 44: 382-384.

Linley, J.R. 1985. Growth and survival of Culicoides melleus larvae (Diptera: Ceratopogonidae) on four prey organisms. J. Med. Ent. 22: 178-189.

Braverman, Y., Linley, J.R., Marcus, R. and Frish, K. 1985. Seasonal survival and vectorial capacity for bluetongue virus of Culicoides spp. (Diptera: Ceratopogonidae) from Israel and a comparison of the parous rate in C. imicola from Israel and Zimbabwe. J. Med. Ent. 22: 476-484.

Linley, J.R. 1985. Biting midges (Diptera: Ceratopogonidae) as vectors of nonviral animal pathogens. J. Med. Ent. 22: 589-599.

Linley, J.R. 1985. Two BASIC programs for statistical analysis of periodicity data, based on the sine-wave function. J. Am. Mosq. Control Assn. 1: 251-253.

Linley, J.R. 1986. Larval behavior. Pp 43-49. in "Bionomics and physiology of Aedes taeniorhynchus and Aedes sollicitans, the salt marsh mosquitos of Florida." Ed. J.K. Nayar. Florida Agricultural Experiment Station, Institution of Food and Agricultural Sciences, Tech. Bull. No. 852, 148 pp.

Linley, J.R. and Braverman, Y. 1986. The lateral abdominal pigmentation in Culicoides variipennis and Culicoides furens (Diptera: Ceratopogonidae): quantitative measurement of its relationship to age and oogenesis. J. Med. Ent. 23: 51-63.

Linley, J.R., Linley, P.A. and Lounibos, L.P. 1986. Light and scanning electron microscopy of the egg of Mansonia titillans (Diptera: Culicidae). J. Med. Ent. 23: 51-63.

Linley, J.R. 1986. The effect of salinity on oviposition and egg hatching in Culicoides variipennis sonorensis (Diptera: Ceratopogonidae). J. Am. Mosq. Control Assoc. 2: 79-82.

Linley, J.R. 1986. Swimming behavior of the larva of Culicoides variipennis (Diptera: Ceratopogonidae) and its relationship to temperature and viscosity. J. Med. Ent. 23: 473-483.

Linley, J.R. 1987. Diel rhythm and lifetime course of oviposition in Toxorhynchites amboinensis (Diptera: Culicidae). J. Med. Ent. 24: 99-105.

Linley, J.R. 1987. Aerial oviposition flight of Toxorhynchites amboinensis (Diptera: Culicidae). J. Med. Ent. 24: 637-650.

Borovsky, D., Linley, J.R. and Kagan, J. 1987. Polycyclic aromatic compounds as phototoxic mosquito larvacides. J. Am. Mosq. Contr. Assn. 3: 246-250.

Linley, J.R., Parsons, R.E. and Winner, R.A. 1987. Evaluation of naled applied as a thermal fog against Culicoides furens (Diptera: Ceratopogonidae). J. Am. Mosq. Contr. Assn. 3: 387-391.

Lounibos, L.P. and Linley, J.R. 1987. A quantitative analysis of underwater oviposition by the mosquito, Mansonia titillans. Physiol. Ent. 12: 435-443.

Linley, J.R. 1988. Note on critical-point drying of mosquito eggs for scanning electron microscopy. J. Med. Ent. 25: 301-302.

Chan, K.L. and Linley, J.R. 1988. Description of Atrichopogon wirthi new species (Diptera: Ceratopogonidae) from leaves of the water lettuce (Pistia stratiotes) in Florida. Fla. Ent. 71: 186-201.

Linley, J.R. 1988. Laboratory experiments on factors affecting oviposition site selection in Toxorhynchites amboinensis (Diptera: Culicidae) with a report on the occurrence of egg cannibalism. Med. Vet. Ent. 2: 271-277.

Linley, J.R., Parsons, R.E. and Winner, R.A. 1988. Evaluation of ULV naled applied simultaneously against caged adult Aedes taeniorhynchus and Culicoides furens. J. Am. Mosq. Contr. Assn. 4: 326-332.

Chan, K.L. and Linley, J.R. 1989. Laboratory studies on the immature stages of Atrichopogon wirthi (Diptera: Ceratopogonidae). Fla. Ent. 72: 85-88.

Linley, J.R. 1989. Egg of Mansonia dyari described and compared with egg of Mansonia titillans (Diptera: Culicidae). J. Med. Ent. 26: 41-45.

Linley, J.R. 1989. Scanning electron microscopy of the eggs of Toxorhynchites rutilus rutilus and Tx. amboinensis (Diptera: Culicidae). J. Med. Ent. 26: 1-9.

Chan, K.L. and Linley, J.R. 1989. A new Florida species of Forcipomyia (Euprojoannisia) (Diptera: Ceratopogonidae) from leaves of the water lettuce, Pistia stratiotes. Fla. Ent. 72: 252-262.

Linley, J.R. and Clark, G.C. 1989. Egg of Aedes (Gymnometopa) mediovittatus (Diptera: Culicidae). J. Med. Ent. 26: 252-255.

Linley, J.R. 1989. Scanning electron microscopy of the egg of Aedes (Protomacleaya) triseriatus (Diptera: Culicidae). J. Med. Ent. 26: 474-478.

Linley, J.R. and Duzak, D. 1989. Egg cannibalism and carnivory among three species of Toxorhynchites (Diptera: Culicidae). J. Am. Mosq. Contr. Assn. 5: 359-362.

Linley, J.R. 1989. Tergal spines of Mansonia titillans and Ma. dyari (Diptera: Culicidae) and their effect on a leaf surface during oviposition. J. Med. Ent. 26: 402-406.

Linley, J.R. 1989. Egg cannibalism in Toxorhynchites r. rutilus (Diptera: Culicidae). J. Fla. Anti-Mosq. Assn. 60: 21-22.

Linley, J.R. 1989. Laboratory tests of the effects of p-cresol and 4-methylcyclohexanol on oviposition by three species of Toxorhynchites mosquitos. Med. Vet. Ent. 3: 347-352.

Linley, J.R. 1989. Comparative fine structure of the eggs of Aedes albopictus, Ae. aegypti and Ae. bahamensis (Diptera: Culicidae). J. Med. Ent. 26: 510-521.

Linley, J.R. and Seabury, F. 1990. Egg of Toxorhynchites splendens described and compared with the egg of Toxorhynchites amboinensis (Diptera: Culicidae). J. Med. Ent. 27: 9-13.

Linley, J.R. 1990. The predatory behavior of Toxorhynchites amboinensis and Tx. brevipalpis larvae (Diptera: Culicidae) in response to subsurface prey. Fla. Ent. 73: 9-51.

Chan, K.L. and Linley, J.R. 1990. The distribution of immature Atrichopogon wirthi (Diptera: Ceratopogonidae) on leaves of the water lettuce Pistia stratiotes. Environ. Ent. 19: 286-292.

Linley, J.R., Lounibos, L.P. and Linley, P.A. 1990. Fine structure of the egg of Trichoprosopon digitatum (Diptera: Culicidae) and its relationship to egg raft formation. J. Med. Ent. 27: 578-585.

Magnon, G.J., Hagan, D.V., Kline, D.L. and J.R. Linley. 1990. Habitat characteristics and phenology of larval Culicoides spp. (Diptera: Ceratopogonidae) from a coastal Georgia salt marsh. Env. Ent. 19 (4): 1068-1074.

Wirth, W.W. and Linley, J.R. 1990. Description of Dasyhelea chani new species (Diptera: Ceratopogonidae) from leaves of the water lettuce (Pistia stratiotes L.) in Florida. Fla. Ent. 73: 274-279.

Linley, J.R. and Chadee, D.D. 1990. Fine structure of the eggs of Psorophora columbiae, Ps. cingulata and Ps. ferox (Diptera: Culicidae). Proc. Ent. Soc. Wash. 92: 497-511.

Linley, J.R. 1990. Scanning electron microscopy of the eggs of Aedes vexans and Aedes infirmatus (Diptera: Culicidae). Proc. Ent. Soc. Wash. 92: 685-693.

Chan, K.L. and Linley, J.R. 1991. Distribution of immature Dasyhelea chani (Diptera: Ceratopogonidae) on leaves of Pistia stratiotes. Ann. Ent. Soc. Am. 84: 61-66.

Linley, J.R. and Chadee, D.D. 1991. The eggs of Haemagogus equinus and Hg. janthinomys (Diptera: Culicidae). J. Med. Ent. 28: 434-445.

Linley, J.R. and Chan, K.L. 1991. Scanning electron microscopy of the eggs of Aedes togoi and Tripteroides bambusa (Diptera: Culicidae). Trop. Biomed. 8: 39-47.

Linley, J.R., Geary, M.J. and Russell, R.C. 1991. The eggs of Aedes funereus, Aedes notoscriptus and Aedes alternans (Diptera: Culicidae). Proc. Ent. Soc. Wash. 93: 592-612.

Linley, J.R. 1991. The egg of Trichoprosopon compressum (Diptera: Culicidae). Mosq. Syst. 23: 161-169.

Linley, J.R., Geary, M.J. and Russell, R.C. 1991. The eggs of Aedes (Finlaya) alboannulatus and Aedes (Finlaya) rubrithorax (Diptera: Culicidae). Mosq. Syst. 23: 132-143.

Linley, J.R., Geary, M.J. and Russell, R.C. 1991. Fine structure of the eggs of Aedes (Ochlerotatus) theobaldi, Ae. (Och.) sagax and Ae. (Och.) procax (Diptera: Culicidae). Mosq. Syst. 23: 144-159.

Linley, J.R., Geary, M.J. and Russell, R.C. 1992. The eggs of Aedes vigilax and Aedes vittiger (Diptera: Culicidae). Proc. Ent. Soc. Wash. 94: 48-58.

Linley, J.R., Geary, M.J. and Russell, R.C. 1992. The eggs of Aedes australis and Aedes comptorhynchus (Diptera: Culicidae). Mosq. Syst. 24: 29-39.

Linley, J.R. 1992. The eggs of Anopheles atropos and Anopheles darlingi (Diptera: Culicidae). Mosq. Syst. 24: 40-50.

Linley, J.R. and Jordan, S. 1992. Effects of ultra-low volume and thermal fog malathion, Scourge and naled applied against caged adult Culicoides furens and Culex quinquefasciatus in open and vegetated terrain. J. Am. Mosq. Control Assoc. 8: 69-76.

Linley, J.R., Abo-Ghalia, A.H., Soliman, B.A. and Mukwaya, L.G. 1993. The eggs of Aedes caspius and Aedes africanus (Diptera: Culicidae). Mosq. Syst. 25: 25-34.

Linley, J.R. and Craig, G.B. 1993. The egg of Aedes hendersoni and a comparison of its structure with the egg of Aedes triseriatus (Diptera: Culicidae). Mosq. Syst. 25: 65-76.

Linley, J.R. and K. Darling. 1993. Search behavior associated with egg cannibalism in Toxorhynchites amboinensis and Toxorhynchites rutilus rutilus (Diptera: Culicidae). J. Med. Ent. 30 (3): 561-570.

Braverman, Y. and J.R. Linley. 1993. Effect of light trap height on catch of Culicoides (Diptera: Ceratopogonidae) in Israel. J. Med. Ent. 30 (6): 1060-1063.

Aussel, J.P. and Linley, J.R. 1993. Activity of Culicoides furens larvae (Diptera: Ceratopogonidae) as related to light and nutritional state. J. Med. Ent. 30 (5): 878-882.

Linley, J.R., Kaiser, P.E., and Cockburn, A.F. 1993. A description and morphometric study of the eggs of species of the Anopheles quadrimaculatus complex (Diptera: Culicidae). Mosq. Syst. 25: 124-147.

Linley, J.R. and Lounibos, L.P. 1993. The eggs of Anopheles (Nyssorhynchus) rangeli and Anopheles (Nys.) dunhami (Diptera: Culicidae). Mosq. Syst. 25: 157-160.

Linley, J.R. and M.J. Turell. 1993. The eggs of Aedes dentatus and Aedes fowleri (Diptera: Culicidae). Proc. Ent. Soc. Wash. 95 (1): 7-16.

Linley, J.R., L.P. Lounibos and J. Conn. 1993. A description and morphometric analysis of the eggs of four South American populations of Anopheles (Nyssorhynchus) aquasalis (Diptera: Culicidae). Mosquito Syst. 25 (3): 198-214.

Aussel, J.P. and J.R. Linley. 1994. Natural food and feeding behavior of Culicoides furens larvae (Diptera: Ceratopogonidae). J. Med. Ent. 31 (1): 90-104.

Linley, J.R. and L.P. Lounibos. 1994. The remarkable egg of Anopheles peryassui (Diptera: Culicidae). Mosquito Syst. 26 (1): 25-34.

Linley, J.R. and M.J. Turell. 1994. Ultrastructural description of the eggs of Aedes mcintoshi and Aedes circumluteolus (Diptera: Culicidae). Mosquito Syst. 26 (1): 1-10.

Linley, J.R. and P.E. Kaiser. 1994. The eggs of Anopheles punctipennis and Anopheles perplexens (Diptera: Culicidae). Mosquito Syst. 26 (2): 43-56.

Linley, J.R. and M.W. Service. 1994. Fine structure of the eggs of Aedes durbanensis, Aedes woodi and Eretmapodites quinquevittatus (Diptera: Culicidae). Mosquito Syst. 26 (2): 59-72.

Linley, J.R., and A.H. Benton and J.F. Day. 1994. Ultrastructure of the eggs of seven flea species (Siphonaptera). J. Med. Ent. 31 (6): 813-827.

Braverman, Y. and J.R. Linley. 1994. Fecundity and proportions of gravid females in populations of the bluetongue vector Culicoides imicola (Diptera: Ceratopogonidae) and several other species in Israel. J. Med. Ent. 31 (6): 838-843.

Linley, J.R. and G.B. Craig. 1994. Morphology of long- and short-day eggs of Aedes atropalpus and Aedes epactius (Diptera: Culicidae). J. Med. Ent. 31 (6): 855-867.

Linley, J.R. 1995. Behavior on approach to surface prey by larvae of Toxorhynchites amboinensis and Toxorhynchites brevipalpis (Diptera: Culicidae). J. Med. Ent. 32 (1): 53-65.

Linley, J.R. and E.G. Milstey. 1995. The eggs of Anopheles mattogrossensis and Chagasia fajardi (Diptera: Culicidae). Mosquito Systematics: In Press

Linley, J.R., H.H. Yap and T.B. Damar. 1995. The eggs of four Malaysian species of Anopheles in the Hyrcanus Group. Mosquito Systematics: In Press